In an Angle’s Class I occlusion, the cusp of which permanent tooth is in contact with the central fossa of the mandibular first molar?

Mesiolingual cusp of the maxillary first molar.

Mesiolingual cusp of the maxillary second molar.

Distolingual cusp of the maxillary second molar.

Distolingual cusp of the maxillary first molar.